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COLUMBUS, Ohio (AP) — She is a mother of three, grandmother of 16, great-grandmother of 12 and great-great-grandmother of three. She recently had surgery to remove a malignant tumor, but doctors say she's doing well. She's Colo, the nation's oldest living gorilla, and she's turning 60 on Thursday at the Columbus Zoo and Aquarium. Colo was the first gorilla in the world born in a zoo and has surpassed the usual life expectancy of captive gorillas by two decades. Her longevity is putting a spotlight on the medical care, nutrition and up-to-date therapeutic techniques that are helping lengthen zoo animals' lives. "Colo just epitomizes the advances that zoos have made, going all the way back to her birth at Columbus," said Dr. Tom Meehan, vice president for veterinary services at Chicago's Brookfield Zoo and veterinary adviser to a national gorilla species survival plan. The changes also mean more animals living with the normal aches and pains of growing older. Today, zoo veterinarians regularly treat animals for heart and kidney disease, arthritis, dental problems and cancer. Colo is one of several elderly gorillas around the country. The oldest known living male gorilla, Ozzie, is 55 years old and lives at the Atlanta Zoo, which has a geriatric gorilla specialty. At Seattle's Woodland Park Zoo, staff members use acupuncture, massage, laser therapy, and heat and joint supplements to help Emma, a 13-year-old rabbit. At the National Zoo in Washington, Shanthi, a 42-year-old Asian elephant with arthritis, receives osteoarthritis therapy and was recently fitted with specially crafted front foot boots to help her feet heal as medications are applied. In Oakland, California, Tiki, a 27-year-old giraffe and one of the oldest in the nation, gets foot care, massage therapy, acupuncture and chiropractic care, along with traditional veterinary medicine. Gao Gao, a 26-year-old male panda at the San Diego Zoo with a heart condition, periodically undergoes cardiac ultrasounds. "Geriatrics is probably one of our most common medical challenges that we face in a zoo situation," said Dr. Keith Hinshaw, director of animal health at the Philadelphia Zoo. "So pretty much anything that you could imagine would happen with an older person is going to happen eventually with any animal." That's up to and including medication: JJ, a 45-year-old orangutan at the Toledo Zoo, is on the human heart medicines carvedilol and Lisinopril, along with pain and orthopedic medications. He also takes Metamucil. Colo, a western lowland gorilla, holds several other records. On her 56th birthday in 2012, she exceeded the record for longest-lived gorilla. On Thursday, she surpasses the median life expectancy for female gorillas in human care (37.5 years) by more than two decades. Other age-defying zoo animals: ___ POLAR BEAR Coldilocks, a 36-year-old polar bear at the Philadelphia Zoo and considered the oldest polar bear in the U.S. The bears' typical lifespan in captivity is 23 years. The zoo says treating her early for kidney disease appears to have helped prolong her life. ___ RHINO Elly, an eastern black rhino at the San Francisco Zoo estimated to be 46 years old, is the oldest of her species in North America. She has had 14 calves, and her offspring have produced 15 grandchildren, 6 great-grandchildren and 1 great-great-grandchild. ___ ELEPHANT Packy, an Asian elephant at the Oregon Zoo, and at 54, the oldest male of his species in North America. The zoo says Packy, born in 1962, became the first elephant to be born in the Western Hemisphere in 44 years. ___ MONKEY Nikko, a 33-year-old snow monkey at the Minnesota Zoo, the oldest male snow monkey in North America. ___ CHIMPANZEE Little Mama, a chimpanzee living at Lion Country Safari in Loxahatchee, Florida, with an estimated age in her late 70s. She takes allergy medicine, iron supplements and omega 3 multivitamins, and has been trained to accept a nebulizer treatment for coughing. ___ TORTOISE Emerson, a Galapagos tortoise at the Toledo Zoo in Ohio, whose age is estimated at about 100. ___ Andrew Welsh-Huggins can be reached on Twitter at https://twitter.com/awhcolumbus. His work can be found at http://bigstory.ap.org/content/andrew-welsh-huggins ___ This story has been corrected to show that Tiki the giraffe is in Oakland, California, not San Francisco.

Happy birthday to Colo: Oldest gorilla in US turns 60